On The Origin of Species by Charles Darwin Chapter 12 Page 6

inhabitants of the eastern and western shores of South America are very distinct, with extremely few shells, crustacea, or echinodermata in common; but Dr. Gunther has recently shown that about thirty per cent of the fishes are the same on the opposite sides of the isthmus of Panama; and this fact has led naturalists to believe that the isthmus was formerly open. Westward of the shores of America, a wide space of open ocean extends, with not an island as a halting-place for emigrants; here we have a barrier of another kind, and as soon as this is passed we meet in the eastern islands of the Pacific with another and totally distinct fauna. So that three marine faunas range northward and southward in parallel lines not far from each other, under corresponding climate; but from being separated from each other by impassable barriers, either of land or open sea,