On The Origin of Species by Charles Darwin Chapter 15 Page 10

always have been a good chance for wide migration by many means. A broken or interrupted range may often be accounted for by the extinction of the species in the intermediate regions. It cannot be denied that we are as yet very ignorant as to the full extent of the various climatical and geographical changes which have affected the earth during modern periods; and such changes will often have facilitated migration. As an example, I have attempted to show how potent has been the influence of the Glacial period on the distribution of the same and of allied species throughout the world.

We are as yet profoundly ignorant of the many occasional means of transport. With respect to distinct species of the same genus, inhabiting distant and isolated regions, as the process of modification has necessarily been slow, all the means