On The Origin of Species by Charles Darwin Chapter 5 Page 12

groups of beetles, elsewhere excessively numerous, which absolutely require the use of their wings, are here almost entirely absent. These several considerations make me believe that the wingless condition of so many Madeira beetles is mainly due to the action of natural selection, combined probably with disuse. For during many successive generations each individual beetle which flew least, either from its wings having been ever so little less perfectly developed or from indolent habit, will have had the best chance of surviving from not being blown out to sea; and, on the other hand, those beetles which most readily took to flight would oftenest have been blown to sea, and thus destroyed.

The insects in Madeira which are not ground-feeders, and which, as certain flower-feeding coleoptera and lepidoptera, must habitually