On The Origin of Species by Charles Darwin Chapter 7 Page 40

quadrupeds, for instance South America, though it is so luxuriant, while South Africa abounds with them to an unparalleled degree. Why this should be so we do not know; nor why the later tertiary periods should have been much more favourable for their existence than the present time.

Whatever the causes may have been, we can see that certain districts and times would have been much more favourable than others for the development of so large a quadruped as the giraffe.

In order that an animal should acquire some structure specially and largely developed, it is almost indispensable that several other parts should be modified and coadapted. Although every part of the body varies slightly, it does not follow that the necessary parts should always vary in the right direction and to the right