On The Origin of Species by Charles Darwin Chapter 7 Page 94

same time drags away some of the pollen-grains. From this simple condition, which differs but little from that of a multitude of common flowers, there are endless gradations — to species in which the pollen-mass terminates in a very short, free caudicle — to others in which the caudicle becomes firmly attached to the viscid matter, with the sterile stigma itself much modified.

In this latter case we have a pollinium in its most highly developed and perfect condition. He who will carefully examine the flowers of orchids for himself will not deny the existence of the above series of gradations — from a mass of pollen-grains merely tied together by threads, with the stigma differing but little from that of the ordinary flowers, to a highly complex pollinium, admirably adapted for transportal by insects; nor