On The Origin of Species by Charles Darwin Chapter 8 Page 82

profitable to the community that a number should have been annually born capable of work, but incapable of procreation, I can see no especial difficulty in this having been effected through natural selection.

But I must pass over this preliminary difficulty. The great difficulty lies in the working ants differing widely from both the males and the fertile females in structure, as in the shape of the thorax, and in being destitute of wings and sometimes of eyes, and in instinct. As far as instinct alone is concerned, the wonderful difference in this respect between the workers and the perfect females would have been better exemplified by the hive-bee. If a working ant or other neuter insect had been an ordinary animal, I should have unhesitatingly assumed that all its characters had been slowly acquired through natural