On The Origin of Species by Charles Darwin Chapter 8 Page 94

workers of a different size and structure; a graduated series having first been formed, as in the case of the driver ant, and then the extreme forms having been produced in greater and greater numbers, through the survival of the parents which generated them, until none with an intermediate structure were produced.

An analogous explanation has been given by Mr. Wallace, of the equally complex case, of certain Malayan butterflies regularly appearing under two or even three distinct female forms; and by Fritz Muller, of certain Brazilian crustaceans likewise appearing under two widely distinct male forms. But this subject need not here be discussed.

I have now explained how, I believe, the wonderful fact of two distinctly defined castes of sterile workers existing in the same nest, both