On The Origin of Species by Charles Darwin Chapter 11 Page 46

beneath the uppermost line may be considered as extinct. The three existing genera, a14, q14, p14, will form a small family; b14 and f14, a closely allied family or subfamily; and o14, i14, m14, a third family. These three families, together with the many extinct genera on the several lines of descent diverging from the parent form (A) will form an order; for all will have inherited something in common from their ancient progenitor.

On the principle of the continued tendency to divergence of character, which was formerly illustrated by this diagram, the more recent any form is the more it will generally differ from its ancient progenitor. Hence, we can understand the rule that the most ancient fossils differ most from existing forms. We must not, however, assume that divergence of character is a necessary