On The Origin of Species by Charles Darwin Chapter 11 Page 65

We ought also to compare the relative proportional numbers, at any two periods, of the high and low classes throughout the world: if, for instance, at the present day fifty thousand kinds of vertebrate animals exist, and if we knew that at some former period only ten thousand kinds existed, we ought to look at this increase in number in the highest class, which implies a great displacement of lower forms, as a decided advance in the organisation of the world. We thus see how hopelessly difficult it is to compare with perfect fairness, under such extremely complex relations, the standard of organisation of the imperfectly-known faunas of successive periods.

We shall appreciate this difficulty more clearly by looking to certain existing faunas and floras. From the extraordinary manner in which European productions have