On The Origin of Species by Charles Darwin Chapter 4 Page 114

(A); the two groups, moreover, are supposed to have gone on diverging in different directions.

The intermediate species, also (and this is a very important consideration), which connected the original species (A) and (I), have all become, except (F), extinct, and have left no descendants. Hence the six new species descended from (I), and the eight descendants from (A), will have to be ranked as very distinct genera, or even as distinct sub-families.

Thus it is, as I believe, that two or more genera are produced by descent with modification, from two or more species of the same genus. And the two or more parent-species are supposed to be descended from some one species of an earlier genus. In our diagram this is indicated by the