On The Origin of Species by Charles Darwin Chapter 4 Page 121

Although few of the most ancient species have left modified descendants, yet, at remote geological periods, the earth may have been almost as well peopled with species of many genera, families, orders and classes, as at the present day.

9. On The Degree To Which Organisation Tends To Advance

Natural selection acts exclusively by the preservation and accumulation of variations, which are beneficial under the organic and inorganic conditions to which each creature is exposed at all periods of life.

The ultimate result is that each creature tends to become more and more improved in relation to its conditions. This improvement inevitably leads to the gradual advancement of the organisation of the greater number of living beings throughout the world.