On The Origin of Species by Charles Darwin Chapter 9 Page 69

we find, and this seems to me of especial importance, that two or three forms of the same species may exist and may differ in no respect whatever, either in structure or in constitution, relatively to external conditions, and yet be sterile when united in certain ways. For we must remember that it is the union of the sexual elements of individuals of the same form, for instance, of two long- styled forms, which results in sterility; while it is the union of the sexual elements proper to two distinct forms which is fertile.

Hence the case appears at first sight exactly the reverse of what occurs, in the ordinary unions of the individuals of the same species and with crosses between distinct species. It is, however, doubtful whether this is really so; but I will not enlarge on this obscure subject.