On The Origin of Species by Charles Darwin Chapter 14 Page 120

supplanted by their successors with the same organ in a more perfect state, and consequently will have become long ago extinct. The wing of the penguin is of high service, acting as a fin; it may, therefore, represent the nascent state of the wing: not that I believe this to be the case; it is more probably a reduced organ, modified for a new function: the wing of the Apteryx, on the other hand, is quite useless, and is truly rudimentary.

Owen considers the simple filamentary limbs of the Lepidosiren as the “beginnings of organs which attain full functional development in higher vertebrates;” but, according to the view lately advocated by Dr. Gunther, they are probably remnants, consisting of the persistent axis of a fin, with the lateral rays or branches aborted. The mammary glands of the Ornithorhynchus may